ALSUP, District Judge:
Jane Crawford appeals her jury convictions for two counts each of wire fraud and interstate transportation of property stolen, converted, or taken through fraud. The jury implicitly found that Crawford took a valuable oil painting from her office at UCLA, intending to deprive the owner of its use, and sold it for profit, knowing that it had been stolen, converted, or taken through fraud. The jury also implicitly found that Crawford (or a co-schemer) had sent and received interstate faxes to advance her scheme to obtain money through false pretenses. The dispositive issue in this appeal is whether sufficient evidence supported the convictions. We conclude that it did, and affirm.
Statement
Frost Flowers, If switch 1889, an oil painting by the influential arts-and-crafts artist Arthur Wesley Dow, hung on the wall of Crawford’s UCLA office in the early 1990s.3 Crawford was UCLA’s Director of Counseling, College of Letters and Sciences. In 1995, she took the painting from her office and, with the help of a middleman named Ken Weaver, secretly sold it.
In May 1999, a federal grand jury indicted Crawford on two counts of wire fraud in violation of 18 U.S.C. § 1343, and two counts of interstate transportation of property converted and obtained through fraud, in violation of 18 U.S.C. § 2314.4 Crawford pleaded not guilty, and a three-[1089]*1089day jury trial began. At the end of the government’s case, and after hearing argument, the court denied Crawford’s Rule 29 motion for acquittal. The jury found Crawford guilty on all four counts.
The parties agree that Arthur Wesley Dow’s widow gave Frost Flowers, along with seven other Dow paintings, to the Arthur Wesley Dow Association in 1928. The government presented evidence that the Association had been affiliated with the University of California, Southern Branch — the precursor to UCLA. In addition, “Los Angeles Normal School” was stenciled on the back of the canvas. Los Angeles Normal School, a teachers’ college in the 1880s, later became part of the University of California, Southern Branch. When the Arthur Wesley Dow Association dissolved, the government asserted, Frost Flowers remained with UCLA (or its predecessor).
Craig Cunningham, a UCLA employee supervised by Crawford, first saw Frost Flowers in UCLA’s Office of the Registrar in Murphy Hall in the 1970s. In 1979, he took the painting to his office at the urging of the Registrar, who was remodeling. When Cunningham found that his office did not have enough wall space, he took the painting home. Frost Flowers hung over his mantel for about ten years. He testified that he had never thought of Frost Floioers as his, but considered it UCLA’s property. He did not hide it from his UCLA supervisors and colleagues when they visited his home, and he told them that it belonged to the University.
In the early 1990s he brought it back to UCLA, placing it in a storage room, and mentioning it to Crawford because he knew that she had admired the painting. She retrieved it from the storage room, and hung it on her office wall, where it remained for several years. During that time, she had conversations with her UCLA colleague Judith Collas in which, Collas testified, Crawford indicated her understanding that UCLA owned the painting. In one such conversation, they spoke about the “irony that UCLA owned such a valuable painting and didn’t know it.”
In 1994, however, Crawford began preparations to sell the painting. The government presented evidence that she called her father’s trust attorney and asked about selling some paintings, including Frost Flowers by Arthur Wesley Dow, that her father had bought, she falsely claimed, while living in Utah years before. At Crawford’s (or her father’s) direction, the attorney wrote a “to whom it may concern” letter on the trust’s behalf, noting that it had retained Kenneth Weaver to secure buyers for the paintings. In March 1995, Crawford called her tax accountant to ask about the tax consequences of selling a painting that had been given to her, she said, by a professor friend at UCLA.
In June 1995, Weaver contracted with Spanierman Gallery to sell Frost Flowers for $200,000. The contract was faxed between California and New York for signatures. Weaver deposited the $200,000 from Spanierman Gallery into an account in his name. He wrote several checks on the account to pay for goods and services for Crawford. He also wrote checks on the account to pay off her debt.
Meanwhile, Crawford’s UCLA colleague Judith Collas noticed that Frost Flowers was no longer in Crawford’s office. Crawford told her that a student at Cal Arts in Valencia was restoring it, under a professor’s supervision. She told Cunningham the same story. In fact, there was no evidence that Crawford had ever sent the painting to be restored, and a Spanierman Gallery representative testified that it arrived in poor condition.
The government called Michael Trenta-lange, UCLA’s Executive Director of External Affairs, to testify about the University’s policies for acquiring and disposing of property. Before his current post, he had served as Executive Director of Information Systems and as Executive Director of Gift Policy and Information Systems.
[1090]*1090When an “affiliated organization” dissolves, Trentalange said, its property remains with UCLA. He testified over Crawford’s objection that UCLA considers an “affiliated organization” to be one that uses the University’s name, facilities, personnel, or equipment in its activities.
He next testified about UCLA’s procedures for disposing of property. Typically a unit head — such as a dean, a director, or a faculty chairperson — decides that a particular asset is no longer needed or wanted. The University then offers the property to its other departments. If no department wants it, the University sells it. Since at least 1981, the University has had “an obligation to get the highest and best return on the sale of property.” When the University decides to dispose of art, it auctions it to the highest bidder. UCLA cannot give property away, Tren-talange testified. Nor, he testified over Crawford’s objection, can it abandon property.
Trentalange further testified that for all items above a certain dollar threshold in value, each UCLA department keeps an inventory, describing the item and listing its location, value, means of acquisition, and means of disposal (if appropriate). An item’s absence from an inventory list, Trentalange testified, does not mean that the University does not claim ownership. If an item were undervalued, for example, it might inadvertently be left off the list.
On cross-examination, Trentalange testified that he had learned of three people who identified the painting as having hung in Murphy Hall since the 1940s. Based on that fact, he executed a declaration in 1998 that UCLA owned Frost Flowers.
Crawford now appeals her conviction, arguing that the district court abused its discretion in admitting much of Trenta-lange’s testimony, and that insufficient evidence of the University’s ownership supported the jury’s verdict.
Analysis
Crawford’s appeal raises two questions.
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ALSUP, District Judge:
Jane Crawford appeals her jury convictions for two counts each of wire fraud and interstate transportation of property stolen, converted, or taken through fraud. The jury implicitly found that Crawford took a valuable oil painting from her office at UCLA, intending to deprive the owner of its use, and sold it for profit, knowing that it had been stolen, converted, or taken through fraud. The jury also implicitly found that Crawford (or a co-schemer) had sent and received interstate faxes to advance her scheme to obtain money through false pretenses. The dispositive issue in this appeal is whether sufficient evidence supported the convictions. We conclude that it did, and affirm.
Statement
Frost Flowers, If switch 1889, an oil painting by the influential arts-and-crafts artist Arthur Wesley Dow, hung on the wall of Crawford’s UCLA office in the early 1990s.3 Crawford was UCLA’s Director of Counseling, College of Letters and Sciences. In 1995, she took the painting from her office and, with the help of a middleman named Ken Weaver, secretly sold it.
In May 1999, a federal grand jury indicted Crawford on two counts of wire fraud in violation of 18 U.S.C. § 1343, and two counts of interstate transportation of property converted and obtained through fraud, in violation of 18 U.S.C. § 2314.4 Crawford pleaded not guilty, and a three-[1089]*1089day jury trial began. At the end of the government’s case, and after hearing argument, the court denied Crawford’s Rule 29 motion for acquittal. The jury found Crawford guilty on all four counts.
The parties agree that Arthur Wesley Dow’s widow gave Frost Flowers, along with seven other Dow paintings, to the Arthur Wesley Dow Association in 1928. The government presented evidence that the Association had been affiliated with the University of California, Southern Branch — the precursor to UCLA. In addition, “Los Angeles Normal School” was stenciled on the back of the canvas. Los Angeles Normal School, a teachers’ college in the 1880s, later became part of the University of California, Southern Branch. When the Arthur Wesley Dow Association dissolved, the government asserted, Frost Flowers remained with UCLA (or its predecessor).
Craig Cunningham, a UCLA employee supervised by Crawford, first saw Frost Flowers in UCLA’s Office of the Registrar in Murphy Hall in the 1970s. In 1979, he took the painting to his office at the urging of the Registrar, who was remodeling. When Cunningham found that his office did not have enough wall space, he took the painting home. Frost Flowers hung over his mantel for about ten years. He testified that he had never thought of Frost Floioers as his, but considered it UCLA’s property. He did not hide it from his UCLA supervisors and colleagues when they visited his home, and he told them that it belonged to the University.
In the early 1990s he brought it back to UCLA, placing it in a storage room, and mentioning it to Crawford because he knew that she had admired the painting. She retrieved it from the storage room, and hung it on her office wall, where it remained for several years. During that time, she had conversations with her UCLA colleague Judith Collas in which, Collas testified, Crawford indicated her understanding that UCLA owned the painting. In one such conversation, they spoke about the “irony that UCLA owned such a valuable painting and didn’t know it.”
In 1994, however, Crawford began preparations to sell the painting. The government presented evidence that she called her father’s trust attorney and asked about selling some paintings, including Frost Flowers by Arthur Wesley Dow, that her father had bought, she falsely claimed, while living in Utah years before. At Crawford’s (or her father’s) direction, the attorney wrote a “to whom it may concern” letter on the trust’s behalf, noting that it had retained Kenneth Weaver to secure buyers for the paintings. In March 1995, Crawford called her tax accountant to ask about the tax consequences of selling a painting that had been given to her, she said, by a professor friend at UCLA.
In June 1995, Weaver contracted with Spanierman Gallery to sell Frost Flowers for $200,000. The contract was faxed between California and New York for signatures. Weaver deposited the $200,000 from Spanierman Gallery into an account in his name. He wrote several checks on the account to pay for goods and services for Crawford. He also wrote checks on the account to pay off her debt.
Meanwhile, Crawford’s UCLA colleague Judith Collas noticed that Frost Flowers was no longer in Crawford’s office. Crawford told her that a student at Cal Arts in Valencia was restoring it, under a professor’s supervision. She told Cunningham the same story. In fact, there was no evidence that Crawford had ever sent the painting to be restored, and a Spanierman Gallery representative testified that it arrived in poor condition.
The government called Michael Trenta-lange, UCLA’s Executive Director of External Affairs, to testify about the University’s policies for acquiring and disposing of property. Before his current post, he had served as Executive Director of Information Systems and as Executive Director of Gift Policy and Information Systems.
[1090]*1090When an “affiliated organization” dissolves, Trentalange said, its property remains with UCLA. He testified over Crawford’s objection that UCLA considers an “affiliated organization” to be one that uses the University’s name, facilities, personnel, or equipment in its activities.
He next testified about UCLA’s procedures for disposing of property. Typically a unit head — such as a dean, a director, or a faculty chairperson — decides that a particular asset is no longer needed or wanted. The University then offers the property to its other departments. If no department wants it, the University sells it. Since at least 1981, the University has had “an obligation to get the highest and best return on the sale of property.” When the University decides to dispose of art, it auctions it to the highest bidder. UCLA cannot give property away, Tren-talange testified. Nor, he testified over Crawford’s objection, can it abandon property.
Trentalange further testified that for all items above a certain dollar threshold in value, each UCLA department keeps an inventory, describing the item and listing its location, value, means of acquisition, and means of disposal (if appropriate). An item’s absence from an inventory list, Trentalange testified, does not mean that the University does not claim ownership. If an item were undervalued, for example, it might inadvertently be left off the list.
On cross-examination, Trentalange testified that he had learned of three people who identified the painting as having hung in Murphy Hall since the 1940s. Based on that fact, he executed a declaration in 1998 that UCLA owned Frost Flowers.
Crawford now appeals her conviction, arguing that the district court abused its discretion in admitting much of Trenta-lange’s testimony, and that insufficient evidence of the University’s ownership supported the jury’s verdict.
Analysis
Crawford’s appeal raises two questions. The first is whether the district court abused its discretion by admitting Trenta-lange’s recounting of the UCLA definition of “affiliated entity,” that the property of an “affiliated entity” stays with the University when the entity dissolves, that UCLA is not authorized to abandon property, and that an asset’s absence from University inventory lists does not indicate that UCLA makes no claim of title to the asset. The second is whether sufficient evidence supported the jury’s implicit findings that Crawford had the requisite mens rea for any of the charged crimes. In particular, Crawford argues that the government did not produce sufficient evidence that UCLA owned Frost Flowers, and therefore did not produce sufficient evidence to prove criminal intent to deprive the rightful owner of its possession.
A. Evidentiary Issues
The Court reviews evidentiary rulings at trial for abuse of discretion. Old Chief v. United States, 519 U.S. 172, 174 n. 1, 117 S.Ct. 644, 136 L.Ed.2d 574 (1997). The Court reviews only for plain error the admission of testimony to which the defendant failed to object at trial. United States v. Hanley, 190 F.3d 1017, 1029 (9th Cir.1999).
A lay witness may testify only to “those opinions or inferences which are (a) rationally based on the perception of the witness and (b) helpful to a clear understanding of the witness’ testimony or the determination of a fact in issue.” Fed. R.Evid. 701. A lay witness may testify as to an ultimate issue of fact, so long as the testimony is otherwise admissible. Fed. R.Evid. 704. The lay witness may not, however, testify as to a legal conclusion, such as the correct interpretation of a contract. Evangelista v. Inlandboatmen’s Union of Pacific, 111 F.2d 1390, 1398 n. 3 (9th Cir.1985).
At trial, Crawford objected that three questions put to Trentalange called for legal conclusions. Crawford alleges that the court abused its discretion in overruling each objection, and that it plainly erred [1091]*1091in admitting two other instances of legal conclusion testimony, even though no objection was raised to the first and the second was elicited on cross-examination, As discussed below, the court did not abuse its discretion or otherwise err, except in allowing Trentalange’s testimony that UCLA could not abandon property,
1. Objections Timely Made During Trial
First, defense counsel objected when the government asked Trentalange to “please explain what an affiliated organization is for the jury.” After the court overruled the objection, Trentalange explained that “We consider an affiliated organization an organization that uses the University’s name, facilities, personnel or equipment in the course of their activities.”
The court did not abuse its discretion by overruling the objection. The question did not call for Trentalange to explain the legal definition of “affiliated organization,” or to conclude that the Arthur Wesley Dow Association was an “affiliated organization.” Nor did Trentalange so testify. He simply told the jury, based on his experience with UCLA’s policies, how UCLA used the term “affiliated organization.”
Second, defense counsel unsuccessfully objected when the government asked Trentalange “can UCLA abandon its property?” Explaining his answer of “no,” Trentalange testified, “Again the same principle that I’ve mentioned before. If it has value, we have an obligation to retain that value, to either keep it or sell it and use those funds for other purposes on campus.”
The court abused its discretion here. “Abandon” is a legal term, and the jury could have read Trentalange’s statement to mean that UCLA cannot, as a matter of law, be found to have “abandoned” property. This interpretation is supported by his reference to his earlier testimony that “[everything the University has belongs to the public essentially; it’s a public institution, and it would be the equivalent of giving away public resources.” If the jury believed that UCLA could not have legally abandoned the painting, the jury would likely conclude that Crawford must have stolen it, regardless of any evidence that UCLA did not claim ownership of it. For the reasons stated below, however, the error was harmless.
Third, defense counsel objected, to no avail, when the government asked “does the fact that a particular item doesn’t appear on an inventory mean that the University is basically saying we don’t own it?” Trentalange answered, “No.” He did not conclude that, in such circumstances, UCLA would or would not own a particular item; he only testified that such a circumstance would not signify that the University intended to forgo ownership. He expressed no opinion as to whether UCLA owned Frost Flowers. Nor did the question ask him to so opine. The court did not abuse its discretion.
2. No Objection Made At Trial
Repeating her legal-conclusion argument, Crawford challenges on appeal the admission of Trentalange’s testimony that the property of an “affiliated organization” belongs to the University and that, when the organization dissolves, its property remains with the University. Crawford did not object to the specific question that elicited this testimony, she did not explicitly make a continuing objection, and she did not move to strike. Nevertheless, she assumes without explanation that the admission of the testimony should be reviewed as though she had. The government notes the lack of an objection, and argues that the admission of the testimony should be reviewed for plain error. The Court agrees with the government that absent a specific objection, an explicit continuing objection, or circumstances that effectively precluded either of the two foregoing options, the admission of evidence should be reviewed for plain error.
The court did not plainly err by admitting the testimony. Trentalange testified as to the policy at UCLA, with which he [1092]*1092was familiar, for handling property of dissolved “affiliated organizations,” a UCLA term. Trentalange did not interpret disputed law or offer an opinion about who owned Frost Flowers.
3. Testimony Elicited on Cross-Examination
Finally, in response to a series of specific questions on cross-examination, Trentalange testified that he had asserted UCLA’s claim of ownership in a sworn 1998 declaration: “I hereby advise the court on behalf of UCLA that UCLA presently claims it is the exclusive owner of the Dow painting and is entitled to sole possession of the Dow painting.” He testified that he made the declaration because he had “learned of ... three employees who identified the painting as having hung in Murphy Hall since the 1940s, for 40 or more years.” The government had not covered this subject on direct examination. In these circumstances, any possible error was invited by defense counsel, who broached the subject of the declaration and asked the questions about its basis. See United States v. Cabrera, 201 F.3d 1243, 1249 (9th Cir.2000) (noting that defense counsel’s own question invited any error). “[A]n error that is caused by the actions of the complaining party will cause reversal only in the most exceptional situation ... [where] reversal is necessary to preserve the integrity of the judicial process or to prevent a miscarriage of justice.” Id. (quoting United States v. Schaff, 948 F.2d 501, 506 (9th Cir.1991)). The circumstances here do not warrant reversal based on the admission of this testimony. As discussed below, any error in its admission was harmless.
B. Sufficiency of the Evidence
Sufficient evidence supports a conviction if the evidence, viewed in the light most favorable to the prosecution, would allow any rational trier of fact to find the essential elements of the crime beyond a reasonable doubt. Jackson v. Virginia, 443 U.S. 307, 319, 99 S.Ct. 2781, 61 L.Ed.2d 560 (1979).
In her sufficiency-of-the-evidence argument, Crawford challenges only the evidence supporting the jury’s implicit finding that she knowingly stole Frost Flowers. If sufficient evidence did not support a finding that UCLA owned the painting, she contends, then sufficient evidence could not have supported the mens rea required by all counts with which she was charged.5 The government disputes both the premise that it had to prove UCLA’s ownership, and the claim that sufficient evidence did not support such a finding.
The government did not need to prove beyond a reasonable doubt that UCLA owned Frost Flowers, only that Crawford knew that she did not.6 The [1093]*1093days of homesteading are over, and the jury may presume that property has a rightful owner, even if the identity of the rightful owner is not immediately known to one who comes upon the property. See Cal. Civ.Code §§ 2080 & 2080.1 (a finder who takes charge of lost property becomes a depository for the owner, and must return the property if the owner is known, or must turn the property over to the police department if unknown).
Even if title to Frost Flowers had never passed to UCLA, Crawford’s actions met the elements of the charges against her. The evidence at trial supported a finding that Crawford took possession of the painting by falsely claiming to her colleagues that she was having it restored, and by falsely telling others that her father had bought the painting years before; that she caused it to be moved over state lines; and that she intended permanently to deprive the true owner—whether that was UCLA, the heir to the Arthur Wesley Dow Association, or some other entity—of its use.
The interstate-transportation statute simply does not require that the transporter know the identity of the true owner. Likewise, the wire-fraud statute requires only that she have participated in a scheme to “obtain money or property by means of false or fraudulent pretenses, representations, or promises” with the intent to defraud. Neither statute requires that Crawford have known her victim’s identity; only that there was proof from which the jury could conclude that she was not the lawful owner of the artwork entitled to dispose of it.
On appeal, Crawford asserts, and this Court is aware, of only one exception to the general rule that things have owners: abandoned property. Under California law, abandonment requires non-use accompanied by unequivocal and decisive acts showing an intent to abandon. See Pacific Gas & Elec. Co. v. Zuckerman, 189 Cal.App.3d 1113, 1145, 234 Cal.Rptr. 630, 650 (1987). Had Crawford proved the elements of abandonment at trial, she could not have been convicted on any of the counts charged, because her sale of the painting would not have deprived the true owner of its use, and she would not have defrauded anyone. But Crawford did not request an instruction on abandonment. Even if she had, sufficient evidence supported a finding that no owner of the painting engaged in any unequivocal or decisive act showing an intent to abandon it. Moreover, sufficient evidence refuted any inference that, whether or not the painting actually had been abandoned, Crawford believed that the painting was abandoned, and thus lacked mens rea.
The relevant evidence instead showed that Crawford did not own the painting, and knew she did not: (1) she first gained custody of the painting by taking it from a UCLA storage room, or so the jury could infer from Cunningham’s testimony that he placed it in the storage room and told Crawford about it, and Collas’s testimony that the painting then turned up on Crawford’s office wall; (2) while Cunningham had possession of the painting, he represented to her and to others that UCLA owned it; (3) Crawford expressed to Collas her understanding that UCLA owned the painting; (4) she lied to UCLA employees about why the painting no longer hung in her office; (5) she lied to her father’s trust lawyer about who owned the painting and how it had been acquired; (6) she lied to her tax accountant about how she had acquired the painting; and (7) she and Weaver deposited the proceeds into an account in his name, not hers, though the money was also for her benefit.
This evidence sufficiently supports jury findings that Crawford did not own Frost Flowers, that she knew she did not own it, and that she intended to deprive the owner, whoever that may have been, of its use. Because the government did not need to prove UCLA’s ownership, Trenta-lange’s challenged testimony would not likely have affected the verdict and any error in its admission was harmless. See United States v. Yin, 935 F.2d 990, 994 (9th Cir.1991) (“A nonconstitutional evi-[1094]*1094dentiary error will be reversed for an abuse of discretion only if the court’s ruling more likely than not affected the verdict.”).
Conclusion
Sufficient evidence supported the jury’s implicit findings that Crawford did not own Frost Flowers, and that she knew she did not own it. We affirm Crawford’s convictions.
AFFIRMED.