[191] BUTTLER, P. J.
Defendant was convicted of unlawful sale of wildlife. ORS 498.022. On appeal, he raises numerous assignments of error. Because we agree with defendant that the state has not established that it has jurisdiction over this offense, we do not address the other assignments.
Defendant is an enrolled member of the Warm Springs Confederated Tribes. He was convicted of selling a deer, which, pursuant to a tribal permit, had been lawfully killed on the Warm Springs Indian Reservation. The sale occurred off the reservation on land ceded by the Confederated Tribes to the United States under an 1855 treaty that expressly reserved to the Indians fishing and hunting rights.1 State law prohibits the sale of any wildlife; so far as the record shows, tribal law does not. Defendant argues that, because he is a member of a tribe that has reserved hunting rights (including the right to sell game) on off-reservation ceded lands, the state has no authority to enforce its wildlife laws against him.2
As a general rule, states have jurisdiction to enforce non-discriminatory laws against Indians off the reservation. Mescalero Apache Tribe v. Jones, 411 US 145, 148-49, 93 S Ct [192]*1921267, 36 L Ed 2d 114 (1973). In addition, states, under their police power, generally have authority to protect and regulate the taking of wildlife within their boundaries.3 See, e.g., Baldwin v. Montana Fish and Game Comm’n, 436 US 371, 391, 98 S Ct 1852, 56 L Ed 2d 354 (1978). However, when there is an applicable federal statute or treaty, the state’s powers are limited by the Supremacy Clause of the United States Constitution. Sohappy v. Smith, supra n 3, 302 F Supp at 905. Accordingly, state regulation of Indian hunting and fishing must yield to Indian rights secured by a federal treaty.
The state argues that the right to hunt, reserved in the treaty, does not include the right to sell that which is taken pursuant to the right. As appealing as that proposition may be, the authorities are to the contrary. In Puyallup Tribe v. Dept. of Game, 391 US 392, 398, 88 S Ct 1725, 20 L Ed 2d 689 (1968) (Puyallup T), a treaty fishing case, the Supreme Court recognized that commercial uses of the fish resource are one aspect of the treaty fishing right, subject to state regulation only when necessary for conservation. In Antoine v. Washington, supra n 2, the Court recognized that the right to hunt also encompasses the right to use the game for commercial purposes. 420 US at 207. Therefore, the state’s contention that sale of game is beyond the reach of the treaty rights and federal decisions is not correct.
The hunting rights in the treaty are not exclusive to the Indians. They are to be shared “in common with citizens.” Federal caselaw has established that, when the rights are held “in common with” other citizens, states may regulate Indian hunting4 on ceded land,5 but only when the regulation is in the [193] interest of conservation, “provided the regulation meets appropriate standards and does not discriminate against the Indians.” Puyallup I, supra, 391 US at 398.6 In Antoine v. Washington, supra n 2, the Supreme Court explained:
“The ‘appropriate standards’ requirement means that the State must demonstrate that its regulation is a reasonable and necessary conservation measure, and that its application to the Indians is necessary in the interest of conservation.” 420 US at 207. (Citations omitted; emphasis in original.)
Therefore, there are three requirements that must be met before a state has the authority to impose its game regulations on treaty hunters in ceded territory. First, the state must show that the regulation is a reasonable and necessary7 conservation measure. Conservation purposes are narrowly circumscribed, encompassing only those that are necessary for the perpetuation of a species, Washington Game Dept. v. Puyallup Tribe, 414 US 44, 48, 94 S Ct 330, 38 L Ed 2d 254 (1973) (Puyallup II), or “necessary to prevent the exercise of [the hunting] right in a manner that will imperil the continued existence of the * * * resource.” Sohappy v. Smith, supra n 3, 302 F Supp at 908.
Second, the state must show that application of the specific regulation to treaty hunters and fishers is necessary in the interest of conservation. In United States v. State of Washington, supra n 2, Judge Boldt explained:
“If alternative means and methods of reasonable and necessary conservation regulation are available, the state cannot lawfully restrict the exercise of off reservation treaty right fishing [or hunting], even if the only alternatives are restriction of fishing [or hunting] by non-treaty fishermen [or hunters], either commercially or otherwise, to the full extent necessary for conservation of [the resource].” 384 F Supp at 342. (Emphasis supplied.)
[194] Finally, the regulation must not discriminate against treaty hunters. That is to say, the state may not subordinate the treaty right to some other state objective or policy. Sohappy v. Smith, supra n 3, 302 F Supp at 908. In Sohapp, Oregon was found to be discriminating against treaty Indians in its fishery regulations because, although the regulations attempted to make an equitable distribution of the anadromous fish resource between sports fishers and commercial fishers, the state had not taken into consideration the rights of treaty fishers to share in the resource.
When the state seeks to enforce a hunting regulation against treaty Indians in ceded areas, the burden is on the state to establish that its regulation is valid in the light of the foregoing requirements. As Judge Boldt said, “[N]o regulation applied to off reservation treaty [hunting] can be valid or enforceable unless and until it has been shown reasonable and necessary to conservation [as defined by federal law].” United States v. State of Washington, supra n 2, 384 F Supp at 342; see also Maison v. Confederated Tribes of Umatilla Indian Res., 314 F2d 169 (9th Cir), cert den 375 US 829 (1963).
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[191] BUTTLER, P. J.
Defendant was convicted of unlawful sale of wildlife. ORS 498.022. On appeal, he raises numerous assignments of error. Because we agree with defendant that the state has not established that it has jurisdiction over this offense, we do not address the other assignments.
Defendant is an enrolled member of the Warm Springs Confederated Tribes. He was convicted of selling a deer, which, pursuant to a tribal permit, had been lawfully killed on the Warm Springs Indian Reservation. The sale occurred off the reservation on land ceded by the Confederated Tribes to the United States under an 1855 treaty that expressly reserved to the Indians fishing and hunting rights.1 State law prohibits the sale of any wildlife; so far as the record shows, tribal law does not. Defendant argues that, because he is a member of a tribe that has reserved hunting rights (including the right to sell game) on off-reservation ceded lands, the state has no authority to enforce its wildlife laws against him.2
As a general rule, states have jurisdiction to enforce non-discriminatory laws against Indians off the reservation. Mescalero Apache Tribe v. Jones, 411 US 145, 148-49, 93 S Ct [192]*1921267, 36 L Ed 2d 114 (1973). In addition, states, under their police power, generally have authority to protect and regulate the taking of wildlife within their boundaries.3 See, e.g., Baldwin v. Montana Fish and Game Comm’n, 436 US 371, 391, 98 S Ct 1852, 56 L Ed 2d 354 (1978). However, when there is an applicable federal statute or treaty, the state’s powers are limited by the Supremacy Clause of the United States Constitution. Sohappy v. Smith, supra n 3, 302 F Supp at 905. Accordingly, state regulation of Indian hunting and fishing must yield to Indian rights secured by a federal treaty.
The state argues that the right to hunt, reserved in the treaty, does not include the right to sell that which is taken pursuant to the right. As appealing as that proposition may be, the authorities are to the contrary. In Puyallup Tribe v. Dept. of Game, 391 US 392, 398, 88 S Ct 1725, 20 L Ed 2d 689 (1968) (Puyallup T), a treaty fishing case, the Supreme Court recognized that commercial uses of the fish resource are one aspect of the treaty fishing right, subject to state regulation only when necessary for conservation. In Antoine v. Washington, supra n 2, the Court recognized that the right to hunt also encompasses the right to use the game for commercial purposes. 420 US at 207. Therefore, the state’s contention that sale of game is beyond the reach of the treaty rights and federal decisions is not correct.
The hunting rights in the treaty are not exclusive to the Indians. They are to be shared “in common with citizens.” Federal caselaw has established that, when the rights are held “in common with” other citizens, states may regulate Indian hunting4 on ceded land,5 but only when the regulation is in the [193] interest of conservation, “provided the regulation meets appropriate standards and does not discriminate against the Indians.” Puyallup I, supra, 391 US at 398.6 In Antoine v. Washington, supra n 2, the Supreme Court explained:
“The ‘appropriate standards’ requirement means that the State must demonstrate that its regulation is a reasonable and necessary conservation measure, and that its application to the Indians is necessary in the interest of conservation.” 420 US at 207. (Citations omitted; emphasis in original.)
Therefore, there are three requirements that must be met before a state has the authority to impose its game regulations on treaty hunters in ceded territory. First, the state must show that the regulation is a reasonable and necessary7 conservation measure. Conservation purposes are narrowly circumscribed, encompassing only those that are necessary for the perpetuation of a species, Washington Game Dept. v. Puyallup Tribe, 414 US 44, 48, 94 S Ct 330, 38 L Ed 2d 254 (1973) (Puyallup II), or “necessary to prevent the exercise of [the hunting] right in a manner that will imperil the continued existence of the * * * resource.” Sohappy v. Smith, supra n 3, 302 F Supp at 908.
Second, the state must show that application of the specific regulation to treaty hunters and fishers is necessary in the interest of conservation. In United States v. State of Washington, supra n 2, Judge Boldt explained:
“If alternative means and methods of reasonable and necessary conservation regulation are available, the state cannot lawfully restrict the exercise of off reservation treaty right fishing [or hunting], even if the only alternatives are restriction of fishing [or hunting] by non-treaty fishermen [or hunters], either commercially or otherwise, to the full extent necessary for conservation of [the resource].” 384 F Supp at 342. (Emphasis supplied.)
[194] Finally, the regulation must not discriminate against treaty hunters. That is to say, the state may not subordinate the treaty right to some other state objective or policy. Sohappy v. Smith, supra n 3, 302 F Supp at 908. In Sohapp, Oregon was found to be discriminating against treaty Indians in its fishery regulations because, although the regulations attempted to make an equitable distribution of the anadromous fish resource between sports fishers and commercial fishers, the state had not taken into consideration the rights of treaty fishers to share in the resource.
When the state seeks to enforce a hunting regulation against treaty Indians in ceded areas, the burden is on the state to establish that its regulation is valid in the light of the foregoing requirements. As Judge Boldt said, “[N]o regulation applied to off reservation treaty [hunting] can be valid or enforceable unless and until it has been shown reasonable and necessary to conservation [as defined by federal law].” United States v. State of Washington, supra n 2, 384 F Supp at 342; see also Maison v. Confederated Tribes of Umatilla Indian Res., 314 F2d 169 (9th Cir), cert den 375 US 829 (1963).
Although the state concedes that it must meet those conditions in order for it to have jurisdiction to regulate Indian treaty hunting rights, it argues that the conservation purpose of the statute prohibiting the sale of wildlife is self-evident and that, therefore, it should not be required to make a showing that its regulations can be validly applied to treaty hunters. We have no doubt that the state’s laws regulating the taking and sale of wildlife have, as one of their goals, the prevention of “the serious depletion of any indigenous species,” ORS 496.012(1), including deer. Their overall policy, however, is much broader:
“It is the policy of the State of Oregon that wildlife shall be managed to provide the optimum recreational and aesthetic benefits for present and future generations of the citizens of this state.” ORS 496.012(1).
Although there is no question but that the state may regulate the taking and selling of wildlife by non-Indians to achieve a wide variety of management or conservation objectives, it does not havé the same latitude in regulating the rights of treaty Indians to take and sell wildlife in ceded territory. See Sohappy v. Smith, supra n 3, 302 F Supp at 908. [195] That proposition was amplified by the Court of Appeals in United States v. State of Washington, 520 F2d 676, 686 (9th Cir 1975):
“The state’s program for management of the state’s fisheries may appear sound and commendable, but the state shares its rights in those fisheries with another party. It may not force treaty Indians to yield their own protected interests in order to promote the welfare of the state’s other citizens. The state must pursue its goals as best it can by regulating its own non-treaty Indian citizens. The state may secure treaty Indians’ compliance with these regulations only by gaining their acquiescence in its goals. Direct regulation of treaty Indian fishing in the interests of conservation is permissible only after the state has proved unable to preserve a run by forbidding the catching of fish by other citizens under its ordinary police power jurisdiction. Antoine v. Washington, 420 U.S. 194, 95 S.Ct. 944, 952, 43 L.Ed.2d 129 (1975).”
It is apparent that the controlling federal law has gone a long way to protect the fishing and hunting treaty rights of Indians. We agree that prohibiting the sale of deer has a rational relation to conservation, because it eliminates an incentive to kill deer. However, as we understand the federal decisions, that is not enough to authorize the state to enforce its regulation against defendant. Here, the tribe permits the on-reservation taking of one deer per member family per month. The state must show, therefore, that prohibiting the sale off the reservation in ceded territory by tribal members of the lawfully taken deer is reasonable and necessary for the perpetuation of the deer population and that restriction of the sale of deer by non-treaty hunters is insufficient to accomplish conservation of the resource.
Here, there is no such evidence, because the trial court agreed with the state’s contention that it need not present evidence to support its jurisdiction over defendant.8
The special concurrence would have us hold that the state may not enforce its prohibition of the sale of game against defendant, regardless of whether it established the requisite conservation purpose, because he killed the deer [196] lawfully on the reservation. It cites no authority for that proposition, and we know of none. However, if that proposition is correct, then an Indian exercising his treaty right to kill deer off the reservation in ceded territory would also be free of any state laws prohibiting the sale of deer for which a conservation purpose is established, because he would have killed it lawfully pursuant to his treaty right.9 That is not, and should not be, the law; to permit the sale of deer under those circumstances vastly expands the market and offers the incentive to kill more deer.
Part of the special concurrence’s confusion arises out of its assertion that it is irrelevant that defendant sold the deer in ceded territory. Defendant’s treaty right to hunt, including the right to sell game, off the reservation extends only to the “unclaimed lands,” which are territories within which Indians formerly claimed the exclusive right to hunt and fish and in which they reserved those rights, under the treaty, in common with others when that territory was ceded to the United States by the treaty. If defendant had not sold the deer in such a place, he would not have been entitled to claim here that he was exercising his treaty right to sell the deer.
In short, the question in any hunting or fishing case in which the event for which a treaty Indian is prosecuted occurred off the reservation in ceded territory is whether the state has shown that the regulation that it wishes to enforce is necessary for conservation under the standards discussed above. If the state establishes the requisite conservation purposes, it may exercise its authority over treaty Indians exercising their treaty rights. The state loses here, not because [197] it may never exercise its authority under these circumstances, but because it failed to establish the necessary conservation purpose of its regulation.
Reversed.